Science Quiz / Paramaters and variables

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Can you pick the Paramaters and variables?

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V (in Pagel's model of evolution)
k (reproductive delays)
V(z) (quantitative genetics)
λ
p(z) (quantitative genetics)
R (Smith Fretwell Model)
α (in Ornstein Uhlenbeck model)
E (quantitative genetics)
S in parasite virulence
δ in parasite virulence
S (quantitative genetics)
z (quantitative genetics)
Po in semelparity/ iteroparity modelling
h2 (quantitative genetics)
Vp (quantitative genetics)
L (reproductive delays)
I in parasite virulence
Hint
β in parasite virulence
R (quantitative genetics)
W(z) (quantitative genetics)
G (quantitative genetics
Va (quantitative genetics)
θ (in Pagel's model of evolution)
F(n) (reproductive delays)
μ (in Ornstein Uhlenbeck model)
r
α in parasite virulence
P (quantitative genetics)
R0
Fs (aka λs)
m (reproductive delays)
P1 in semelparity/ iteroparity modelling
S^n (reproductive delays)
B (reproductive delays)

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