Most career HRs having never led league

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Can you name the Most HRs having never led league?

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Career home runsPlayerRank on HR list
569
521
509
475
473
465
449
431
428*
427
Career home runsPlayerRank on HR list
426
410*
409*
399
396
384
383*
379
377
377

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Created May 5, 2010ReportNominate
Tags:career, having, home, hrs, league, never, player, run